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Messengers who might go before riders at the track (7)

Ginge I'm struggling with the wordplay on your @18 - could you oblige please?

Hi both tyke51 (much appreciated) & kt17.
The difference between the letters "P" & "R" is "the downward stroke"- the tail.
(P)unners - Rams as under pun² Chambers with the surface alluding to male sheep => (R)unners.

Thank you Ginge - fiendish!

PS
and the best of luck to you also tyke51.

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