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With so many options for the letters migrating north and south - most top and bottom row cells had at least two and many three possible occupants - the endgame looked like being a nightmare. But actually qait resolved fairly easily. My way in was on the bottom row and remembering where Kcit is based. Thanks to Kcit for what looked a formidable challenge but which quickly made sense.

I’m not sure how “qait” came to be. “It”, obviously.

I didn't get round to this until earlier today. I agree the endgame looked daunting at first sight but I hazarded a lucky guess at the top row and everything then fell neatly into place in the bottom row.