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My first crack for a while, as I'd ground my teeth down on previous examples - this one I suspect would be an easy romp for regular solvers.
I've cracked the theme, and ID'd the missing element, but can't see where the capital is in 20 down, nor why it seems doubled up - any suggestions?

I think the 'capital' just stands for ... 'Upper Case'/UC. I don't know why it seems doubled up - I didn't finish off the crossword.

PS. If you'd given the clue, other people might have been able to help - I'm not very good at parsing answers!

The full clue , with added latter, is:
(S)horn put up as capital on rodent.

.UC.TUCO

Oops, added letter

Tuco-Tuco

The tuco-tuco is a neotropical rodent in the family Ctenomyidae. The tuco-tucos belong to the only living genus of the Ctenomyidae family, ''Ctenomys'', but they include approximately 60 different species. The common name, "tuco-tuco" comes from the "tuc-tuc" sound they make while they dig their

Not sure if that is any help as I don't attempt the Listener

Hi jazzy (et al), I think it might be the alternative spelling, tocutuco, required here (although the 4th letter isn't a crosser).

Tucutuco, sorry.

Thanks, malone and ginge.
Finally returned to it after polishing off today's Grauniad Prize, and it just clicked into place.

The way it works is "(s)horn put up" = TUC + "as" = UT + "capital" = UC + "on" = O. I know that "ut" = "as" is obscure, but it's in Chambers. I've only ever come across it meaning "as" in Latin.